Phylum Arthropoda von Siebold, 1848
Class Trilobita Walch, 1771
Order Proetida Fortey and Owens 1975
Superfamily Proetoidea Hawle and Corda 1847
Family Phillipsiidae Oehlert, 1886

Both genera of trilobites found in the Late Pennsylvanian rocks of Armstrong County belong to the family Phillipsiidae. They are both small compared to all extinct trilobites. Researchers have documented the shrinking of Trilobite species until their extinction at the end of the Permian during the most extensive mass extinction.

Their occurrence is rare, with about a dozen examples recovered over three years of collection. The tail, known as the pygidium, is found more often than the thorax or cephalon (the head). In both locations, perhaps the chiton exoskeleton disarticulated after death, making recovery of complete specimens rare.

The Brush Creek and Pine Creek limestone in this part of Armstrong County contains only two genera–Ameura missouriensis and Ditomopyge scitula. The genus Ameura is rare and can attain lengths up to 7 cm, whereas the smaller Ditomopyge is the most common and maxes out at 3 cm (Babcock, 1996, p. 101). They appear similar at first glance, yet the relative height and width of the axial lobe is shorter and narrower in Ameura. Both disappeared from the fossil record before the Ames limestone (Brezinski et al., 1989) in local rocks, but both remain in the midcontinent.

Brezinski et al. (1989) studied the morphological variation of Ditomopyge scitula across the Pottsville, Allegheny, and Conemaugh Groups. The average pygidial width dropped by 2 mm, the pygidial length by 2.5 mm, and the average number of pleural ribs and axial rings reduced. While there is no confirmed reason, all trilobites continued to decrease in average size until their extinction.

Trilobites from Armstrong County, PA.
Fig. 1.—Trilobites from Armstrong County, PA. A, C, D, pygidiums of Ditomopyge scitula from the Pine Creek limestone at SL 6445, A, CG-0386, C, CG-0384, D, CG-0385. B, E, Ameura missouriensis from the Brush Creek limestone at SL 6533, B, a free cheek/genial spine, CG-0094 E, pygidium, CG-0425. Scale bar = 5 mm.
Trilobites from Armstrong County, PA.
Fig. 2.—Trilobites from Armstrong County, PA. A–D, pygidiums from the family Phillipsiidae, Brush Creek limestone at SL 6533, A, Ameura missouriensis, CG-0593, B, CG-0175, C, CG-0096, D, CG-0174. E–F, Ditomopyge scitula from the Pine Creek limestone at SL 6533, E, CG-0591 F, CG-0592. Scale bar = 5 mm.

References

  • Brezinski, D.K., Sturgeon, M.T., Hoare, R.D., 1989. Pennsylvanian Trilobites of Ohio, Report on Investigations No. 142
  • Hawle, I. & Corda, J.C., 1847. Prodrom einer Monographie der böhmischen Trilobiten. J.G. Calvesche Buchhandlung, Prague, 176 pp
  • Oehlert, D. P., 1886. Étude sur quelques trilobites du groupe de Proetidae. Bulletine Society Étude Scientifique d’Angers, 15, 121–143.
  • Shimer, H. W, Shrock, R. R., 1944. Index Fossils of North America, P. 637, 645, 648
  • Walch, J. E. I., 1771. Die Naturgeschichte der Versteinerungen, zur Erläuterung der Knorrischen Sammlung von Merkwurdigkeiten der Natur, Volume 4, Part 3. (J.P. Felßecker: Nürnberg) 303 pp